139. (32103-!-item-!-188;#058&006640)
Taste buds are onion-shaped structures with between 50 and 100 taste cells, each of them has fingerlike projections poking through the opening located at the top of the taste bud called the taste pore.
(A) them has fingerlike projections poking through the opening located at the top of the taste bud called the taste pore
(B) them that have fingerlike projections that poke through an opening located at the top of the taste bud, which they call the taste pore
(C) which has fingerlike projections poking through the opening, called the taste pore, located at the top of the taste bud
(D) which having fingerlike projections that poke through an opening, which is called the taste pore, located at the top of the taste bud
(E) which have fingerlike projections that are poking through an opening located at the top of the taste bud called the taste pore
答案是C
想請問一下each of which是否為該子句之主格?如此一來有S也有V(has)的結構與前面主句間並無連接詞,會不會有run-on的問題呢?